2026 Land Rover Defender vs 2025 Toyota Land Cruiser

This two-row midsize SUV is designed for traveling almost anywhere at almost any time. It shares its platform with the rugged Toyota 4Runner SUV and Tacoma pickup truck, and is equipped with a turbocharged hybrid powertrain better at supplying power than efficiency. Additionally, it is more affordable than it has been in the past, but still commands a premium over its primary rivals. The 2025 Land Cruiser is almost unchanged following its 2024 redesign, which introduced the current 250-series model (known as the Land Cruiser Prado overseas) to the United States.

Verdict: There is a lot to like about the Land Cruiser, especially if you don’t care about fuel economy. Our test vehicle averaged 15.7 mpg, falling well short of expectations. That thirst for fuel easily ranks as the worst thing about this SUV. Also, we recommend upgrading from the base 1958 trim level.

As the rest of the lineup has trended toward luxury, the Land Rover Defender has stayed true to the brand’s roots of tough off-roaders ideal for the military and farmers alike. That’s mainly because Land Rover put off updates for decades, but a redesigned Defender that’s more passenger car than farm implement arrived for 2020, bringing the nameplate back to the U.S. Changes for 2026 include subtle styling changes, a larger touchscreen, and new color and accessory options.

Verdict: The Defender offers an unmatched combination of off-road capability and everyday civility.