2026 Toyota 4Runner vs 2025 Land Rover Defender

The 2025 Defender lineup includes the 90, 110, and 130. The 90 is the short and stubby two-door, the 110 is the standard four-door, and the 130 is the extended four-door. They’re available with turbocharged four-cylinder and six-cylinder engines, as well as a turbocharged or supercharged V8. They’re all equipped for off-roading right out of the box, but you can add more tech for more challenging situations. The big news for this model year is the high-performance 2025 Land Rover Defender OCTA.

Verdict: There is a Land Rover Defender for almost anyone, and any of them can take you farther into the wilderness than you expect. They’re rugged but refined, comfortable yet capable, and plenty of fun to drive. But the Defender comes at a cost, both on the window sticker and at the pump.

Shoppers looking for an old-school SUV should be careful what they wish for, because the 4Runner makes compromises in passenger comfort and interior space to make it one of the better off-roaders on the market. It’s offered as a hybrid, but fuel economy is never as good as a typical crossover SUVs, which will be more comfortable and more affordable. That makes mainstream crossovers better choices for most, but for those who want adventure while carrying passengers, the 4Runner makes sense.

Verdict: The 4Runner’s closest rivals are the Jeep Wrangler and the Ford Bronco. Compared to them, the 4Runner can carry more passengers and offers better fuel economy thanks to an available hybrid powertrain. If adventure is a priority, then the 4Runner is a great choice, but those who want a five- or seven-passenger SUV for daily family duty would be better off with a crossover.