Unlike in a Toyota Tundra, which has the same 381-horsepower 5.7-liter V8 engine that's bolted into the Land Cruiser, you can barely hear the engine when this SUV is accelerating. The roar is hushed to nearly imperceptible levels, with just enough of its characteristic note filtering into the cabin to remind you why the trip computer's average fuel economy number reads less than 15 mpg. We averaged 14.7 mpg on our testing loop, beating the EPA rating of 14 mpg in combined driving.
An eight-speed automatic transmission funnels the motive force to all four wheels through a full-time four-wheel-drive system with a two-speed transfer case. With 90% of the engine's 401 pound-feet of torque available at just 2,200 rpm, first gear delivers a bunch of unexpected thrust. You might want to engage the 'ECT Second' feature using the button on the center console, which starts the SUV in second gear for much smoother launches.
The Land Cruiser's legendary off-road capability is rooted in more than just this component set. For example, there's a Torsen limited-slip differential that automatically distributes power based on available traction. The normal split sends 60% of the power to the rear wheels, but drivers can lock it into an even 50:50 front-to-rear split when that's desirable.
Standard mud-and-snow tires wrap around the Land Cruiser's 18-inch aluminum wheels. Toyota also includes a full-size spare tire, just in case you run into unexpected trouble while traveling far off the grid.
Toyota's brake-based Active Traction Control limits wheelspin, while hill start assist and downhill traction control help to ensure safe forward progress on steep grades. Multi-Terrain Select traction assist is also standard, providing settings including Rock, Loose Rock, Rock & Dirt, Mud & Sand, and Mogul.
Crawl Control is low-speed cruise control for use on challenging terrain, while Off-Road Turn Assist works in conjunction with Crawl Control to tighten the SUV's turning radius. A Multi-Terrain Monitor adds multiple camera views and an inclinometer so that drivers can more easily monitor the situation.
Toyota slides a sophisticated double-wishbone front and four-link rear suspension under the Land Cruiser, and the standard Kinetic Dynamic Suspension System (KDSS) disconnects the sway bar to ensure maximum wheel articulation whenever necessary. Naturally, a bevy of skid plates comes standard to protect the Land Cruiser's underthings.
By now, you're assuming this SUV can cruise over any type of land, and it sure feels that way. We chucked some rutted and rumpled trails at the Land Cruiser, and it seemingly chuckled at our weak attempt to challenge its capabilities.
On pavement, everything about the way this SUV drives is either muffled, numbed, or lubricated to deliver the refinement necessary to justify the price tag. You don't hear or feel much of anything, aside from some occasional discombobulation in the ride quality. You'll need to work the steering wheel more than you might like when parking this remarkably maneuverable beast, but otherwise, the Land Cruiser drives like a Lexus and handles like the nearly 3-ton leviathan it is.
If you're wondering what you're paying for when you buy a Toyota Land Cruiser, much of the cost is associated with its go-anywhere and conquer-anything hardware. And if you're wondering what this SUV can tow, it's rated at 8,100 pounds.